M. A. Altabet, Variations in nitrogen isotopic composition between sinking and suspended particles: implications for nitrogen cycling and particle transformation in the open ocean, Deep Sea Research Part A. Oceanographic Research Papers, vol.35, issue.4, pp.535-554, 1988.

A. Aminot and R. Kerouel, Dosage automatique des nutriments dans les eaux marines: méthodes en flux continu, 2007.

F. Azam and F. Malfatti, Microbial structuring of marine ecosystems, Nature Reviews Microbiology, vol.5, issue.10, pp.782-791, 2007.

A. Bandyopadhyay, T. Elvitigala, E. Welsh, J. Stöckel, M. Liberton et al., Novel Metabolic Attributes of the Genus Cyanothece, Comprising a Group of Unicellular Nitrogen-Fixing Cyanobacteria, mBio, vol.2, issue.5, pp.214-225, 2011.

M. Benavides, N. Agawin, J. Arístegui, J. Peene, and L. J. Stal, Dissolved organic nitrogen and carbon release by a marine unicellular diazotrophic cyanobacterium, Aquatic Microbial Ecology, vol.69, issue.1, pp.69-80, 2013.
URL : https://hal.archives-ouvertes.fr/hal-00999281

I. Berman-frank, D. Spungin, E. Rahav, F. Van-wambeke, K. Turk-kubo et al., Dynamics of transparent exopolymer particles (TEP) during the VAHINE mesocosm experiment in the New Caledonia lagoon, Biogeosciences Discuss, 2016.

H. Berthelot, S. Bonnet, M. Camps, O. Grosso, and T. Moutin, Assessment of the dinitrogen released as ammonium and dissolved organic nitrogen by unicellular and filamentous marine diazotrophic cyanobacteria grown in culture, Frontiers in Marine Science, vol.2, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01790796

H. Berthelot, T. Moutin, S. L'helguen, K. Leblanc, S. Hélias et al., Dinitrogen fixation and dissolved organic nitrogen fueled primary production and particulate export during the VAHINE mesocosm experiment (New Caledonia lagoon), Biogeosciences, vol.12, issue.13, pp.4099-4112, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01231737

H. Berthelot, T. Moutin, S. L'helguen, K. Leblanc, S. Hélias et al., Dinitrogen fixation and dissolved organic nitrogen fueled primary production and particulate export during the VAHINE mesocosm experiment (New Caledonia lagoon), Biogeosciences, vol.12, issue.13, pp.4099-4112, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01231737

I. C. Biegala and P. Raimbault, High abundance of diazotrophic picocyanobacteria (< 3 µm) in a Southwest Pacific coral lagoon, Aquat. Microb. Ecol, vol.51, pp.45-53, 2008.
URL : https://hal.archives-ouvertes.fr/hal-00727031

D. Bombar, C. D. Taylor, S. T. Wilson, J. C. Robidart, A. Rabines et al., Contrasted geographical distribution of N 2 fixation rates and nifH phylotypes in the Coral and Solomon Seas (South-Western Pacific) during austral winter conditions, Diazotroph derived nitrogen supports diatoms growth in the South West Pacific: a quantitative study using nanoSIMS, vol.37, pp.1874-1892, 2015.

S. Bonnet, T. Moutin, M. Rodier, J. M. Grisoni, F. Louis et al., Introduction to the project VAHINE: VAriability of vertical and tropHIc transfer of diazotroph derived N in the south wEst Pacific, Biogeosciences Discuss, 2016.

R. S. Braman and S. A. Hendrix, Nanogram nitrite and nitrate determination in environmental and biological materials by vanadium (III) reduction with chemiluminescence detection, Anal. Chem, vol.61, pp.2715-2718, 1989.

K. L. Casciotti, D. M. Sigman, M. G. Hastings, J. K. Böhlke, and A. Hilkert, Measurement of the Oxygen Isotopic Composition of Nitrate in Seawater and Freshwater Using the Denitrifier Method, Analytical Chemistry, vol.74, issue.19, pp.4905-4912, 2002.

M. J. Church, B. D. Jenkins, D. M. Karl, and J. P. Zehr, Vertical distributions of nitrogen-fixing phylotypes at Stn Aloha in the oligotrophic North Pacific Ocean, Aquatic Microbial Ecology, vol.38, pp.3-14, 2005.

R. Dabundo, M. F. Lehmann, L. Treibergs, C. R. Tobias, M. A. Altabet et al., The Contamination of Commercial 15N2 Gas Stocks with 15N?Labeled Nitrate and Ammonium and Consequences for Nitrogen Fixation Measurements, PLoS ONE, vol.9, issue.10, p.e110335, 2014.

J. E. Dore, R. M. Letelier, M. J. Church, R. Lukas, K. et al., Summer phytoplankton blooms in the oligotrophic North Pacific Subtropical Gyre: Historical perspective and recent observations, Prog. Oceanogr, vol.76, pp.2-38, 2008.

S. Duhamel, K. M. Björkman, and D. M. Karl, Light dependence of phosphorus uptake by microorganisms in the subtropical North and South Pacific Ocean, Aquatic Microbial Ecology, vol.67, issue.3, pp.225-238, 2012.

C. Dupouy, J. Neveux, A. Subramaniam, M. R. Mulholland, J. P. Montoya et al., Satellite captures trichodesmium blooms in the southwestern tropical Pacific, vol.81, pp.13-16, 2000.

S. T. Dyhrman, P. D. Chappell, S. T. Haley, J. W. Moffett, E. D. Orchard et al., Phosphonate utilization by the globally important marine diazotroph Trichodesmium, Nature, vol.439, issue.7072, pp.68-71, 2006.

R. W. Eppley, F. M. Reid, and J. D. Strickland, Estimates of phytoplankton crop size, growth rate, and primary production, Bull. Scripps Inst. Oceanogr, vol.17, pp.33-42, 1970.

P. G. Falkowski, Evolution of the nitrogen cycle and its influence on the biological sequestration of CO2 in the ocean, Nature, vol.387, issue.6630, pp.272-275, 1997.

S. E. Fawcett, M. W. Lomas, J. R. Casey, B. B. Ward, and D. M. Sigman, Assimilation of upwelled nitrate by small eukaryotes in the Sargasso Sea, Nature Geoscience, vol.4, issue.10, pp.717-722, 2011.

A. A. Fong, D. M. Karl, R. Lukas, R. M. Letelier, J. P. Zehr et al., Influence of the Amazon River plume on distributions of free-living and symbiotic cyanobacteria in the western tropical north Atlantic Ocean, Limnol. Oceanogr, vol.2, pp.517-532, 2007.

R. A. Foster, S. Sztejrenszus, and M. M. Kuypers, Measuring carbon and N 2 fixation in field populations of colonial and free?living unicellular cyanobacteria using nanometer?scale secondary ion mass spectrometry 1, Journal of Phycology, vol.49, issue.3, pp.502-516, 2013.


J. A. Fuhrman, Marine viruses and their biogeochemical and ecological effects, Nature, vol.399, issue.6736, pp.541-548, 1999.

R. Fukuda, H. Ogawa, T. Nagata, and I. Koike, Direct Determination of Carbon and Nitrogen Contents of Natural Bacterial Assemblages in Marine Environments, Applied and Environmental Microbiology, vol.64, issue.9, pp.3352-3358, 1998.

N. Garcia, P. Raimbault, E. Gouze, and V. Sandroni, Fixation de diazote et production primaire en Méditerranée occidentale, Comptes Rendus Biologies, vol.329, issue.9, pp.742-750, 2006.

N. Garcia, P. Raimbault, and V. Sandroni, Seasonal nitrogen fixation and primary production in the Southwest Pacific: nanoplankton diazotrophy and transfer of nitrogen to picoplankton organisms, Marine Ecology Progress Series, vol.343, pp.25-33, 2007.

J. M. Gasol, U. L. Zweifel, F. Peters, J. A. Fuhrman, A. Hagstrom et al., Biogeochemical fluxes and fate of diazotroph derived nitrogen in the food web after a phosphate enrichment: Modeling of the VAHINE mesocosms experiment, Biogeosciences Discuss, vol.65, pp.4475-4483, 1999.

P. M. Glibert and D. Bronk, Release of dissolved organic nitrogen by marine diazotrophic cyanobacteria, Trichodesmium spp, Appl. Environ. Microb, vol.60, pp.3996-4000, 1994.

N. L. Goebel, K. A. Turk, K. M. Achilles, R. Paerl, I. Hewson et al., Abundance and distribution of major groups of diazotrophic cyanobacteria and their potential contribution to N2 fixation in the tropical Atlantic Ocean, Environmental Microbiology, vol.12, issue.12, pp.3272-3289, 2010.

M. N. Grabowski, M. J. Church, and D. M. Karl, Nitrogen fixation rates and controls at Stn ALOHA, Aquatic Microbial Ecology, vol.52, pp.175-183, 2008.

T. Großkopf, W. Mohr, T. Baustian, H. Schunck, D. Gill et al., Doubling of marine dinitrogen-fixation rates based on direct measurements, Nature, vol.488, issue.7411, pp.361-364, 2012.

N. ;. Gruber, M. Oguz, and T. , The Dynamics of the Marine Nitrogen Cycle and its Influence on Atmospheric CO2 Variations, The Ocean Carbon Cycle and Climate, pp.97-148, 2004.

G. R. Hasle, The inverted microscope, Phytoplankton Manual, 1978.

C. Höckelmann, P. G. Becher, S. H. Reuß, and F. Jüttner, Sesquiterpenes of the Geosmin-Producing Cyanobacterium Calothrix PCC 7507 and their Toxicity to Invertebrates, Zeitschrift für Naturforschung C, vol.64, issue.1-2, pp.49-55, 2009.

B. P. Hunt, S. Bonnet, H. Berthelot, B. J. Conroy, R. A. Foster et al., Contribution and pathways of diazotroph derived nitrogen to zooplankton during the VAHINE mesocosm experiment in the oligotrophic New Caledonia lagoon, Biogeosciences Discuss, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01443627

T. M. Kana, C. Darkangelo, M. D. Hunt, J. B. Oldham, G. E. Bennett et al., A membrane inlet mass spectrometer for rapid high precision determination of N 2 , O 2 , and Ar in environmental water samples, Anal. Chem, vol.66, pp.4166-4170, 1994.

D. M. Karl, R. M. Letelier, L. Tupas, J. Dore, J. Christian et al., The role of nitrogen fixation in biogeochemical cycling in the subtropical North Pacific Ocean, Nature, vol.388, issue.6642, pp.533-538, 1997.

D. M. Karl, M. J. Church, J. E. Dore, R. M. Letelier, and C. Mahaffey, Predictable and efficient carbon sequestration in the North Pacific Ocean supported by symbiotic nitrogen fixation, Proceedings of the National Academy of Sciences, vol.109, issue.6, pp.1842-1849, 2012.

A. N. Knapp, J. Dekaezemacker, S. Bonnet, J. A. Sohm, and D. G. Capone, Sensitivity of Trichodesmium erythraeum and Crocosphaera watsonii abundance and N<sub>2 fixation rates to varying NO<sub>3<sup>? and PO<sub>4<sup>3? concentrations in batch cultures, Aquatic Microbial Ecology, vol.66, issue.3, pp.223-236, 2012.

A. N. Knapp, D. M. Sigman, and F. Lipschultz, N isotopic composition of dissolved organic nitrogen and nitrate at the Bermuda Atlantic Time-series Study site, Global Biogeochemical Cycles, vol.19, issue.1, pp.1-15, 2005.

A. N. Knapp, S. E. Fawcett, A. Martínez-garcia, N. Leblond, T. Moutin et al., Nitrogen isotopic evidence for a shift from nitrate- to diazotroph-fueled export production in VAHINE mesocosm experiments, Biogeosciences Discussions, vol.12, issue.23, pp.19901-19939, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01443230

K. Leblanc, J. Arístegui, L. Armand, P. Assmy, B. Beker et al., A global diatom database -abundance, biovolume and biomass in the world ocean, Earth Syst. Sci. Data, vol.4, pp.149-165, 2012.
URL : https://hal.archives-ouvertes.fr/hal-00756934

K. Leblanc, J. Arístegui, L. Armand, P. Assmy, B. Beker et al., A global diatom database ? abundance, biovolume and biomass in the world ocean, Earth System Science Data, vol.4, issue.1, pp.149-165, 2012.
URL : https://hal.archives-ouvertes.fr/hal-00756934

O. Levitan, G. Rosenberg, I. ?etlík, E. ?etlíkova, J. Gtigel et al., Elevated CO 2 enhances nitrogen fixation and growth in the marine cyanobacterium Trichodesmium, Glob. Change Biol, vol.13, pp.1-8, 2007.

Y. Luo, S. C. Doney, L. A. Anderson, M. Benavides, I. Berman-frank et al., Database of diazotrophs in global ocean: abundance, biomass and nitrogen fixation rates, Earth System Science Data, vol.4, issue.1, pp.47-73, 2012.

D. Marie, F. Partensky, N. Simon, L. Guillou, D. Vaulot et al., Flow cytometry analysis of marine picoplankton, in: living colors: protocols in flow cytometry and cell sorting, 2000.

T. B. Meador, L. I. Aluwihare, and C. Mahaffey, Isotopic heterogeneity and cycling of organic nitrogen in the oligotrophic ocean, Limnology and Oceanography, vol.52, issue.3, pp.934-947, 2007.

M. M. Mills, C. Ridame, M. Davey, J. La-roche, and R. J. Geider, Iron and phosphorus co-limit nitrogen fixation in the eastern tropical North Atlantic, Nature, vol.429, issue.6989, pp.292-294, 2004.
URL : https://hal.archives-ouvertes.fr/hal-00166836

W. Mohr, T. Großkopf, D. W. Wallace, and J. Laroche, Methodological Underestimation of Oceanic Nitrogen Fixation Rates, PLoS ONE, vol.5, issue.9, p.e12583, 2010.

P. H. Moisander, E. Rantajärvi, M. Huttunen, and K. Kononen, Phytoplankton community in relation to salinity fronts at the entrance to the Gulf of Finland, Baltic Sea, Ophelia, vol.46, issue.3, pp.187-203, 1997.

P. H. Moisander, R. A. Beinart, M. Voss, and J. P. Zehr, Diversity and abundance of diazotrophic microorganisms in the South China Sea during intermonsoon, The ISME Journal, vol.2, issue.9, pp.954-967, 2008.

P. H. Moisander, R. A. Beinart, I. Hewson, A. E. White, K. S. Johnson et al., Unicellular Cyanobacterial Distributions Broaden the Oceanic N2 Fixation Domain, Science, vol.327, issue.5972, pp.1512-1514, 2010.

P. H. Moisander, R. A. Beinart, I. Hewson, A. E. White, K. S. Johnson et al., Unicellular Cyanobacterial Distributions Broaden the Oceanic N2 Fixation Domain, Science, vol.327, issue.5972, pp.1512-1514, 2010.

J. P. Montoya, M. Voss, P. Kahler, and D. G. Capone, A Simple, High-Precision, High-Sensitivity Tracer Assay for N(inf2) Fixation., Applied and environmental microbiology, vol.62, issue.3, pp.986-993, 1996.

C. M. Moore, M. M. Mills, K. R. Arrigo, I. Berman-frank, L. Bopp et al., Processes and patterns of oceanic nutrient limitation, Nature Geoscience, vol.6, issue.9, pp.701-710, 2013.

T. Moutin, T. F. Thingstad, F. Van-wambeke, D. Marie, G. Slawyk et al., Does competition for nanomolar phosphate supply explain the predominance of the cyanobacterium Synechococcus ?, Limnology and Oceanography, vol.47, issue.5, pp.1562-1567, 2002.
URL : https://hal.archives-ouvertes.fr/hal-01780324

T. Moutin, B. Van-den-b, B. Beker, C. Dupouy, P. Rimmelin et al., Phosphate availability controls Trichodesmium spp. biomass in the SW Pacific Ocean, Marine Ecology Progress Series, vol.297, pp.15-21, 2005.

M. R. Mulholland, D. A. Bronk, and D. G. Capone, Dinitrogen fixation and release of ammonium and dissolved organic nitrogen by Trichodesmium IMS101, Aquatic Microbial Ecology, vol.37, pp.85-94, 2004.

M. R. Mulholland, P. W. Bernhardt, C. A. Heil, D. A. Bronk, and J. M. O&apos;neil, Nitrogen fixation and release of fixed nitrogen by Trichodesmium spp. in the Gulf of Mexico, Limnology and Oceanography, vol.51, issue.4, pp.1762-1776, 2006.

D. M. Nelson, P. Tréguer, M. A. Brzezinski, A. Leynaert, and B. Quéguiner, Production and dissolution of biogenic silica in the ocean: Revised global estimates, comparison with regional data and relationship to biogenic sedimentation, Global Biogeochemical Cycles, vol.9, issue.3, pp.359-372, 1995.

J. O&apos;neil, M. R. Roman, E. J. Carpenter, D. G. Capone, and J. G. Rueter, Marine Pelagic Cyanobacteria: Trichodesmium and other Diazotrophs, Marine Pelagic Cyanobacteria: Trichodesmium and other Diazotrophs, 1992.

K. M. Orcutt, F. Lipschultz, K. Gundersen, R. Arimoto, A. F. Michaels et al., A seasonal study of the significance of N2 fixation by Trichodesmium spp. at the Bermuda Atlantic Time-series Study (BATS) site, Deep Sea Research Part II: Topical Studies in Oceanography, vol.48, issue.8-9, pp.1583-1608, 2001.

L. Polerecky, B. Adam, J. Milucka, N. Musat, T. Vagner et al., Look@NanoSIMS - a tool for the analysis of nanoSIMS data in environmental microbiology, Environmental Microbiology, vol.14, issue.4, pp.1009-1023, 2012.

M. Pujo-pay and P. Raimbault, improvement of the wet-oxidation procedure for simultaneous determination of particulate organic nitrogen and phosphorus collected on filters, Marine Ecology Progress Series, vol.105, pp.203-207, 1994.

H. Qi, T. B. Coplen, H. Geilmann, W. A. Brand, and J. K. Böhlke, Two new organic reference materials for?13C and?15N measurements and a new value for the?13C of NBS 22 oil, Rapid Communications in Mass Spectrometry, vol.17, issue.22, pp.2483-2487, 2003.

E. Rahav, B. Herut, A. Levi, M. R. Mulholland, and I. Berman-frank, Springtime contribution of dinitrogen fixation to primary production across the Mediterranean Sea, Ocean Science, vol.9, issue.3, pp.489-498, 2013.

O. Raveh, N. David, G. Rilov, and E. Rahav, The Temporal Dynamics of Coastal Phytoplankton and Bacterioplankton in the Eastern Mediterranean Sea, PLOS ONE, vol.10, issue.10, p.e0140690, 2015.

A. Redfield, On the proportions of organic derivatives in sea water and their relation to the composition of plankton, James Johnstone Memorial, pp.177-192, 1934.

M. Rodier and R. Le-borgne, Population dynamics and environmental conditions affecting Trichodesmium spp. (filamentous cyanobacteria) blooms in the south?west lagoon of New Caledonia, Journal of Experimental Marine Biology and Ecology, vol.358, issue.1, pp.20-32, 2008.
URL : https://hal.archives-ouvertes.fr/ird-00288736

M. Rodier and R. Le-borgne, Population and trophic dynamics of Trichodesmium thiebautii in the SE lagoon of New Caledonia. Comparison with T. erythraeum in the SW lagoon, Marine Pollution Bulletin, vol.61, issue.7-12, pp.349-359, 2010.

R. Scharek, M. Latasa, D. M. Karl, and R. R. Bidigare, Temporal variations in diatom abundance and downward vertical flux in the oligotrophic North Pacific gyre, Deep Sea Research Part I: Oceanographic Research Papers, vol.46, issue.6, pp.1051-1075, 1999.

R. Scharek, L. M. Tupas, and D. M. Karl, Diatom fluxes to the deep sea in the oligotrophic North Pacific gyre at Station ALOHA, Marine Ecology Progress Series, vol.182, pp.55-67, 1999.

T. Shiozaki, T. Nagata, M. Ijichi, and K. Furuya, Nitrogen fixation and the diazotroph community in the temperate coastal region of the northwestern North Pacific, Biogeosciences, vol.12, issue.15, pp.4751-4764, 2015.

D. M. Sigman, K. L. Casciotti, M. Andreani, C. Barford, M. Galanter et al., A bacterial method for the nitrogen isotopic analysis of nitrate in seawater and freshwater, Anal. Chem, vol.73, pp.4145-4153, 2001.

T. J. Smayda, From phytoplankton to biomass. 6. Phytoplankton manual, Monographs on oceanographic methodology, 1978.

J. A. Sohm and D. G. Capone, Phosphorus dynamics of the tropical and subtropical north Atlantic: Trichodesmium spp. versus bulk plankton, Marine Ecology Progress Series, vol.317, pp.21-28, 2006.

M. Staal, F. J. Meysman, and L. J. Stal, Temperature excludes N2-fixing heterocystous cyanobacteria in the tropical oceans, Nature, vol.425, issue.6957, pp.504-507, 2003.

A. Subramaniam, P. L. Yager, E. J. Carpenter, C. Mahaffey, K. M. Bjorkman et al., Amazon River enhances diazotrophy and carbon sequestration in the tropical North Atlantic Ocean, Proceedings of the National Academy of Sciences, vol.105, issue.30, pp.10460-10465, 2008.

J. Sun and D. Liu, Geometric models for calculating cell biovolume and surface area for phytoplankton, Journal of Plankton Research, vol.25, issue.11, pp.1331-1346, 2003.
URL : https://hal.archives-ouvertes.fr/hal-01663379

A. Thompson, B. J. Carter, K. Turk-kubo, F. Malfatti, F. Azam et al., Genetic diversity of the unicellular nitrogen-fixing cyanobacteria UCYN-A and its prymnesiophyte host, Environmental Microbiology, vol.16, issue.10, pp.3238-3249, 2014.

H. J. Tripp, S. R. Bench, K. A. Turk, R. A. Foster, B. A. Desany et al., Metabolic streamlining in an open-ocean nitrogen-fixing cyanobacterium, Nature, vol.464, issue.7285, pp.90-94, 2010.

K. A. Turk-kubo, I. E. Frank, M. E. Hogan, A. Desnues, S. Bonnet et al., Diazotroph community succession during the VAHINE mesocosm experiment (New Caledonia lagoon), Biogeosciences, vol.12, issue.24, pp.7435-7452, 2015.

F. Van-wambeke, U. Pfreundt, A. Barani, H. Berthelot, T. Moutin et al., Heterotrophic bacterial production and metabolic balance during the VAHINE mesocosm experiment in the New Caledonia lagoon, Biogeosciences Discussions, vol.12, issue.23, pp.19861-19900, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01381792

P. G. Verity, C. Y. Robertson, C. R. Tronzo, M. G. Andrews, J. R. Nelson et al., Relationships between cell volume and the carbon and nitrogen content of marine photosynthetic nanoplankton, Limnology and Oceanography, vol.37, issue.7, pp.1434-1446, 1992.

D. Vincent, G. Slawyk, S. L&apos;helguen, G. Sarthou, M. Gallinari et al., Net and gross incorporation of nitrogen by marine copepods fed on 15N-labelled diatoms: Methodology and trophic studies, Journal of Experimental Marine Biology and Ecology, vol.352, issue.2, pp.295-305, 2007.
URL : https://hal.archives-ouvertes.fr/hal-00482361

A. E. Walsby, The Gas Vesicles and Buoyancy of Trichodesmium, Marine Pelagic Cyanobacteria: Trichodesmium and other Diazotrophs, vol.362, pp.141-161, 1992.

A. E. White, R. A. Foster, C. R. Benitez-nelson, P. Masqué, E. Verdeny et al., Nitrogen fixation in the Gulf of California and the Eastern Tropical North Pacific, Progress in Oceanography, vol.109, pp.1-17, 2013.

S. T. Wilson, D. Böttjer, M. J. Church, and D. M. Karl, Comparative Assessment of Nitrogen Fixation Methodologies, Conducted in the Oligotrophic North Pacific Ocean, Applied and Environmental Microbiology, vol.78, issue.18, pp.6516-6523, 2012.

C. S. Yentsch and D. A. Phinney, Spectral fluorescence: an ataxonomic tool for studying the structure of phytoplankton populations, Journal of Plankton Research, vol.7, issue.5, pp.617-632, 1985.

J. P. Zehr, S. R. Bench, B. J. Carter, I. Hewson, F. Niazi et al., Globally Distributed Uncultivated Oceanic N2-Fixing Cyanobacteria Lack Oxygenic Photosystem II, Science, vol.322, issue.5904, pp.1110-1112, 2008.