, Please note that this is an author-produced PDF of an article accepted for publication following peer review. The definitive publisher-authenticated version is available on the publisher Web site

I. Quéau, C. Mingant, M. Riobé, L. Valérian, L. Roy et al., for their technical help. We thank Élodie Fleury for providing oysters placed in the field and Gary H. Wikfors for assistance with english editing and relevant comments. Field data (algal concentrations, PST contents in algae and oysters) were measured within the framework of the project "DAOULEX

, This work is part of a PhD project supported by the "Laboratoire d'Exellence" LabexMer (ANR-10-LABX-19) and co-founded by a grant from the French government under the program "Investissement d'Avenir, This study was carried out with the financial support of the French National Research Agency (ANR), 2013.

B. Aalvik and K. Framstad, Assay and detoxification experiments with Mytilotoxin in mussels (Mythus edulis L.) from Nordåsstraumen, Western Norway, Sarsia, vol.66, issue.2, pp.143-146, 1979.

M. Alunno-bruscia, Y. Bourlès, D. Maurer, S. Robert, J. Mazurié et al., A single bio-energetics growth and reproduction model for the oyster Crassostrea gigas in six Atlantic ecosystems, J. Sea Res, vol.66, issue.4, pp.340-348, 2011.

D. Anderson, D. Kulis, J. Sullivan, and S. Hall, Toxin composition variations in one isolate of the dinoflagellate Alexandrium fundyense, Toxicon, vol.28, issue.8, pp.885-893, 1990.

D. M. Anderson, Toxic algal blooms and red tides : a global perspective, pp.11-16, 1989.

D. M. Anderson, P. M. Glibert, and J. M. Burkholder, Harmful algal blooms and eutrophication : nutrient sources, composition, and consequences, Estuaries, vol.25, issue.4, pp.704-726, 2002.

G. Arzul, M. Seguel, L. Guzman, and E. Erard-le-denn, Comparison of allelopathic properties in three toxic Alexandrium species, J. Exp. Mar. Biol. Ecol, vol.232, issue.2, pp.285-295, 1999.

L. Barillé, J. Prou, M. Héral, and S. Bourgrier, No influence of food quality, but ration-dependent retention efficiencies in the Japanese oyster Crassostrea gigas, J. Exp. Mar. Biol. Ecol, vol.171, issue.1, pp.91-106, 1993.

C. Béchemin, D. Grzebyk, F. Hachame, C. Hummert, and S. Y. Maestrini, Effect of different nitrogen/phosphorus nutrient ratios on the toxin content in Alexandrium minutum, Aquat. Microb. Ecol, vol.20, issue.2, pp.157-165, 1999.

I. Bernard, G. De-kermoysan, and S. Pouvreau, Effect of phytoplankton and temperature on the reproduction of the Pacific oyster Crassostrea gigas : investigation through DEB theory, J. Sea Res, vol.66, issue.4, pp.349-360, 2011.

J. Blanco, A. Moroño, J. Franco, and M. Reyero, PSP detoxification kinetics in the mussel Mytilus galloprovincialis. one-and two-compartment models and the effect of some environmental variables, Mar. Ecol. Prog. Ser, pp.165-175, 1997.

J. Blanco, M. I. Reyero, and J. Franco, Kinetics of accumulation and transformation of paralytic shellfish toxins in the blue mussel Mytilus galloprovincialis, Toxicon, vol.42, issue.7, pp.777-784, 2003.

X. Bodiguel, O. Maury, C. Mellon-duval, F. Roupsard, A. Le-guellec et al., A dynamic and mechanistic model of PCB bioaccumulation in the European hake (Merluccius merluccius), J. Sea Res, vol.62, issue.2, pp.124-134, 2009.

E. Borcier, R. Morvezen, P. Boudry, P. Miner, G. Charrier et al., Effects of bioactive extracellular compounds and paralytic shellfish toxins produced by Alexandrium minutum on growth and behaviour of juvenile great scallops Pecten maximus, Aquat. Toxicol, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01483131

S. Bougrier, P. Geairon, J. Deslous-paoli, C. Bacher, and G. Jonquières, Allometric relationships and effects of temperature on clearance and oxygen consumption rates of Crassostrea gigas (Thunberg), Aquaculture, vol.134, issue.1, pp.143-154, 1995.

F. Boullot, C. Fabioux, H. Hegaret, P. Soudant, P. Boudry et al., Assessment of saxitoxin sensitivity of nerves isolated from the Pacific oyster, Crassostrea gigas, exposed to Alexandrium minutum, Toxicon, vol.149, p.93, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02334360

Y. Bourlès, M. Alunno-bruscia, S. Pouvreau, G. Tollu, D. Leguay et al., Modelling growth and reproduction of the Pacific oyster Crassostrea gigas : advances in the oyster-DEB model through application to a coastal pond, J. Sea Res, vol.62, issue.2, pp.62-71, 2009.

G. Boyer, J. Sullivan, R. Andersen, P. Harrison, and F. Taylor, Effects of nutrient limitation on toxin production and composition in the marine dinoflagellate Protogonyaulax tamarensis, Mar. Biol, vol.96, issue.1, pp.123-128, 1987.

V. Bricelj, M. Greene, J. Lee, and A. Cembella, Growth of the blue mussel Mytilus edulis on toxic Alexandrium fundyense and effects of gut passage on dinoflagellate cells, Toxic Phytoplankton Blooms in the Sea, pp.371-376, 1993.

V. Bricelj, J. Lee, and A. Cembella, Influence of dinoflagellate cell toxicity on uptake and loss of paralytic shellfish toxins in the northern quahog Mercenaria mercenaria, Mar. Ecol. Prog. Ser, vol.74, pp.33-46, 1991.

V. Bricelj, J. Lee, A. Cembella, and D. Anderson, Uptake kinetics of paralytic shellfish toxins from the dinoflagellate Alexandrium fundyense in the mussel Mytilus edulis, Mar. Ecol. Prog. Ser, vol.63, issue.2, pp.177-188, 1990.

V. M. Bricelj and S. E. Shumway, Paralytic shellfish toxins in bivalve molluscs : occurrence, transfer kinetics, and biotransformation, Rev. Fish. Sci, vol.6, issue.4, pp.315-383, 1998.

S. Casas and C. Bacher, Modelling trace metal (Hg and Pb) bioaccumulation in the mediterranean mussel, Mytilus galloprovincialis, applied to environmental monitoring, J. Sea Res, vol.56, issue.2, pp.168-181, 2006.

J. Castrec, P. Soudant, L. Payton, D. Tran, P. Miner et al., Bioactive extracellular compounds produced by the dinoflagellate Alexandrium minutum are highly detrimental for oysters, Aquat. Toxicol, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02324608

A. Cembella, Ecophysiology and metabolism of paralytic shellfish toxins in marine microalgae, NATO-Advanced Study Institute Series, vol.41, pp.381-404, 1998.

A. D. Cembella, S. E. Shumway, and R. Larocque, Sequestering and putative biotransformation of paralytic shellfish toxins by the sea scallop Placopecten magellanicus : seasonal and spatial scales in natural populations, J. Exp. Mar. Biol. Ecol, vol.180, issue.1, pp.1-22, 1994.

A. Chapelle, M. Le-gac, C. Labry, R. Siano, J. Quere et al., The Bay of Brest (France), a new risky site for toxic Alexandrium minutum blooms and PSP shellfish contamination, Harmful Algae News, vol.51, pp.4-5, 2015.

H. Chebib, A. Cembella, and P. Anderson, Differential paralytic shellfish toxin accumulation and detoxification kinetics in transplanted populations of Mytilus edulis exposed to natural blooms of Alexandrium excavatum, 1993.

C. Y. Chen and H. N. Chou, Accumulation and depuration of paralytic shellfish poisoning toxins by purple clam Hiatula rostrata Lighttoot, Toxicon, vol.39, issue.7, pp.1029-1034, 2001.

H. N. Chou, Y. M. Chen, and C. Y. Chen, Variety of PSP toxins in four culture strains of Alexandrium minutum collected from southern Taiwan, Toxicon, vol.43, issue.3, pp.337-340, 2004.

J. J. Dorantes-aranda, K. Campbell, A. Bradbury, C. T. Elliott, D. T. Harwood et al., Comparative performance of four immunological test kits for the detection of Paralytic Shellfish Toxins in Tasmanian shellfish, Toxicon, vol.125, pp.110-119, 2017.

, Marine biotoxins in shellfish-saxitoxin group, EFSA journal, vol.7, issue.4, p.1019, 2009.

M. Eichinger, V. Loizeau, F. Roupsard, A. Le-guellec, and C. Bacher, Modelling growth and bioaccumulation of Polychlorinated biphenyls in common sole (Solea solea), J. Sea Res, vol.64, issue.3, pp.373-385, 2010.

E. Erard-le-denn, M. Morlaix, and J. Dao, Effects of Gyrodinium cf. aureolum on Pecten maximus (post larvae, juveniles and adults), pp.132-136, 1990.

E. P. Espinosa, R. M. Cerrato, G. H. Wikfors, and B. Allam, Modeling food choice in the two suspension-feeding bivalves, Crassostrea virginica and Mytilus edulis, Mar. Biol, vol.163, issue.2, p.40, 2016.

C. Fabioux, Y. Sulistiyani, H. Haberkorn, H. Hégaret, Z. Amzil et al., Exposure to toxic Alexandrium minutum activates the detoxifying and antioxidant systems in gills of the oyster Crassostrea gigas, Harmful Algae, vol.48, pp.55-62, 2015.
URL : https://hal.archives-ouvertes.fr/hal-02556213

S. E. Ford, V. M. Bricelj, C. Lambert, and C. Paillard, Deleterious effects of a nonpst bioactive compound (s) from Alexandrium tamarense on bivalve hemocytes, Mar. Biol, vol.154, issue.2, pp.241-253, 2008.
URL : https://hal.archives-ouvertes.fr/hal-00461303

M. Gourault, S. Petton, Y. Thomas, L. Pecquerie, G. M. Marques et al., Modeling reproductive traits of an invasive bivalve species under contrasting climate scenarios from 1960 to 2100, Journal of Sea Research, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02412677

C. Guallar, C. Bacher, and A. Chapelle, Global and local factors driving the phenology of Alexandrium minutum (Halim) blooms and its toxicity, Harmful Algae, vol.67, pp.44-60, 2017.

M. Guéguen, Modélisation de la détoxication de mollusques bivalves contenant des phycotoxines paralysantes ou diarrhéiques, 2009.

M. Guéguen, M. Bardouil, R. Baron, P. Lassus, P. Truquet et al., Detoxification of Pacific oyster Crassostrea gigas fed on diets of Skeletonema costatum with and without silt, following PSP contamination by Alexandrium minutum, Aquat. Living Resour, vol.21, issue.1, pp.13-20, 2008.

M. Guéguen, R. Baron, M. Bardouil, P. Truquet, H. Haberkorn et al., Modelling of paralytic shellfish toxin biotransformations in the course of Crassostrea gigas detoxification kinetics, Ecol. Model, vol.222, issue.18, pp.3394-3402, 2011.

M. Guéguen, P. Lassus, M. Laabir, M. Bardouil, R. Baron et al., Gut passage times in two bivalve molluscs fed toxic microalgae :Alexandrium minutum, A. catenella and Pseudo-nitzschia calliantha, Aquat. Living Resour, vol.21, issue.1, pp.21-29, 2008.

H. Haberkorn, C. Lambert, N. Le-goïc, M. Guéguen, J. Moal et al., Effects of Alexandrium minutum exposure upon physiological and hematological variables of diploid and triploid oysters, Crassostrea gigas, Aquat. Toxicol, vol.97, issue.2, pp.96-108, 2010.
URL : https://hal.archives-ouvertes.fr/hal-00589357

H. Haberkorn, C. Lambert, N. Le-goïc, J. Moal, M. Suquet et al., Effects of Alexandrium minutum exposure on nutrition-related processes and reproductive output in oysters Crassostrea gigas, Harmful Algae, vol.9, issue.5, pp.427-439, 2010.
URL : https://hal.archives-ouvertes.fr/hal-00589386

G. M. Hallegraeff, A review of harmful algal blooms and their apparent global increase, Phycologia, vol.32, issue.2, pp.79-99, 1993.

G. M. Hallegraeff, D. M. Anderson, A. D. Cembella, and H. O. Enevoldsen, Manual on harmful marine microalgae, 2003.

H. Hégaret, K. B. Brokordt, C. F. Gaymer, K. B. Lohrmann, C. García et al., Effects of the toxic dinoflagellate Alexandrium catenella on histopathogical and escape responses of the Northern scallop Argopecten purpuratus, Harmful Algae, vol.18, pp.74-83, 2012.

H. Hégaret, S. E. Shumway, G. H. Wikfors, S. Pate, and J. M. Burkholder, Potential transport of harmful algae via relocation of bivalve molluscs, Mar. Ecol. Prog. Ser, vol.361, pp.169-179, 2008.

G. Honsell, R. Poletti, M. Pompei, L. Sidari, A. Milandri et al., Alexandrium minutum Halim and PSP contamination in the northern Adriatic Sea (Mediterranean sea). Harmful Toxic Algal Blooms, pp.77-80, 1996.

J. W. Hurst and E. S. Gilfillan, Paralytic shellfish poisoning in Maine. Department of Marine Resources, 1977.

D. F. Hwang and Y. H. Lu, Influence of environmental and nutritional factors on growth, toxicity, and toxin profile of dinoflagellate Alexandrium minutum, Toxicon, vol.38, issue.11, pp.1491-1503, 2000.

T. Jager, A. Barsi, N. T. Hamda, B. T. Martin, E. I. Zimmer et al., Dynamic energy budgets in population ecotoxicology : Applications and outlook, Ecol. Model, vol.280, pp.140-147, 2014.
URL : https://hal.archives-ouvertes.fr/hal-01210211

T. Jager and E. I. Zimmer, Simplified dynamic energy budget model for analysing ecotoxicity data, Ecol. Model, vol.225, pp.74-81, 2012.

S. Kooijman, The synthesizing unit as model for the stoichiometric fusion and branching of metabolic fluxes, Bioph. Chem, vol.73, issue.1, pp.179-188, 1998.

S. Kooijman and J. Bedaux, Analysis of toxicity tests on fish growth, Wat. Res, vol.30, issue.7, pp.1633-1644, 1996.

S. Kooijman and J. Metz, On the dynamics of chemically stressed populations : the deduction of population consequences from effects on individuals, Ecotoxicol. Environ. Saf, vol.8, issue.3, pp.254-274, 1984.

S. A. Kooijman, Pseudo-faeces production in bivalves, J. Sea Res, vol.56, pp.103-106, 2006.

S. A. Kooijman, Dynamic energy and mass budgets in biological systems, 2000.

S. A. Kooijman, Dynamic energy budget theory for metabolic organisation, 2010.

R. W. Kwong, W. Wang, P. K. Lam, and K. Peter, The uptake, distribution and elimination of paralytic shellfish toxins in mussels and fish exposed to toxic dinoflagellates, Aquat. Toxicol, vol.80, issue.1, pp.82-91, 2006.

M. Laabir, Z. Amzil, P. Lassus, E. Masseret, Y. Tapilatu et al., Viability, growth and toxicity of Alexandrium catenella and Alexandrium minutum (Dinophyceae) following ingestion and gut passage in the oyster Crassostrea gigas, Aquat. Living Resour, vol.20, issue.1, pp.51-57, 2007.

M. Lassudrie, P. Soudant, J. Nicolas, C. Fabioux, C. Lambert et al., Interaction between toxic dinoflagellate Alexandrium catenella exposure and disease associated with herpesvirus OsHV-1µvar in Pacific oyster spat Crassostrea gigas, Harmful Algae, vol.45, pp.53-61, 2015.

M. Lassudrie, G. H. Wikfors, I. Sunila, J. H. Alix, M. S. Dixon et al., Physiological and pathological changes in the eastern oyster Crassostrea virginica infested with the trematode Bucephalus sp. and exposed to the toxic dinoflagellate Alexandrium fundyense, J. Invertebr. Pathol, vol.126, pp.51-63, 2015.
URL : https://hal.archives-ouvertes.fr/hal-02558203

P. Lassus, Z. Amzil, R. Baron, V. Séchet, L. Barillé et al., Modelling the accumulation of PSP toxins in Thau lagoon oysters (Crassostrea gigas) from trials using mixed cultures of Alexandrium catenella and Thalassiosira weissflogii, Aquat. Living Resour, vol.20, issue.01, pp.59-67, 2007.

P. Lassus, M. Bardouil, R. Baron, J. Berard, P. Masselin et al., Improving detoxification efficiency of PSP-contaminated oysters (Crassostrea gigas Thunberg), 2005.

P. Lassus, M. Bardouil, P. Massselin, M. Naviner, and P. Truquet, Comparative efficiencies of different non-toxic microalgal diets in detoxification of PSP-contaminated oysters (Crassostrea gigas Thunberg), J. Nat. Toxins, vol.9, issue.1, pp.1-12, 2000.

P. Lassus, R. Baron, P. Garen, P. Truquet, P. Masselin et al., Paralytic shellfish poison outbreaks in the Penze estuary : Environmental factors affecting toxin uptake in the oyster, Crassostrea gigas, Aquat. Living Resour, vol.17, issue.2, pp.207-214, 2004.

R. Lavaud, J. Flye-sainte-marie, F. Jean, A. Emmery, Ø. Strand et al., Feeding and energetics of the great scallop, Pecten maximus, through a DEB model, J. Sea Res, vol.94, pp.5-18, 2014.
URL : https://hal.archives-ouvertes.fr/hal-00917710

A. Lelong, H. Haberkorn, N. Le-goïc, H. Hégaret, and P. Soudant, A new insight into allelopathic effects of Alexandrium minutum on photosynthesis and respiration of the diatom Chaetoceros neogracile revealed by photosynthetic-performance analysis and flow cytometry, Microb. Ecol, vol.62, issue.4, pp.919-930, 2011.
URL : https://hal.archives-ouvertes.fr/hal-00643865

S. C. Leong, A. Murata, Y. Nagashima, and S. Taguchi, Variability in toxicity of the dinoflagellateAlexandrium tamarense in response to different nitrogen sources and concentrations, Toxicon, vol.43, issue.4, pp.407-415, 2004.

S. Li and W. Wang, Radiotracer studies on the feeding of two marine bivalves on the toxic and nontoxic dinoflagellate Alexandrium tamarense, J. Exp. Mar. Biol. Ecol, vol.263, issue.1, pp.65-75, 2001.

S. Li, W. Wang, and D. P. Hsieh, Effects of toxic dinoflagellate Alexandrium tamarense on the energy budgets and growth of two marine bivalves, Mar. Environ. Res, vol.53, issue.2, pp.145-160, 2002.

M. Long, K. Tallec, P. Soudant, C. Lambert, F. Le-grand et al., A rapid quantitative fluorescence-based bioassay to study allelochemical interactions from Alexandrium minutum, Environ. Poll, 2018.
URL : https://hal.archives-ouvertes.fr/hal-02130252

M. Luckenbach, K. Sellner, S. Shumway, and K. Greene, Effects of two bloom-forming dinoflagellates, Prorocentrum minimum and Gyrodinium uncatenum, on the growth and survival of the eastern oyster, Crassostrea virginica (Gmelin 1791), J. Shellfish Res, vol.12, issue.2, pp.411-415, 1993.

H. Ma, B. Krock, U. Tillmann, A. Muck, N. Wielsch et al., Isolation of activity and partial characterization of large non-proteinaceous lytic allelochemicals produced by the marine dinoflagellate Alexandrium tamarense, Harmful Algae, vol.11, pp.65-72, 2011.

L. L. Mafra, V. M. Bricelj, and J. E. Ward, Mechanisms contributing to low domoic acid uptake by oysters feeding on Pseudonitzschia cells. II. Selective rejection, Aquat. Biol, vol.6, pp.213-226, 2009.

A. Moroño, J. Franco, M. Miranda, M. I. Reyero, and J. Blanco, The effect of mussel size, temperature, seston volume, food quality and volume-specific toxin concentration on the uptake rate of PSP toxins by mussels (Mytilus galloprovincialis Lmk), J. Exp. Mar. Biol. Ecol, vol.257, issue.1, pp.117-132, 2001.

E. Navarro, J. Iglesias, and M. Ortega, Natural sediment as a food source for the cockle Cerastoderma edule (L.) : effect of 25 variable particle concentration on feeding, digestion and the scope for growth, J. Exp. Mar. Biol. Ecol, vol.156, issue.1, pp.69-87, 1992.

J. M. Navarro and A. M. Contreras, An integrative response by Mytilus chilensis to the toxic dinoflagellate Alexandrium catenella, Mar. Biol, vol.157, issue.9, pp.1967-1974, 2010.

M. Nielsen and T. Strømgren, Shell growth response of mussels (Mytilus edulis) exposed to toxic microalgae, Mar. Biol, vol.108, issue.2, pp.263-267, 1991.

Y. Oshima, Chemical and enzymatic transformation of paralytic shellfish toxins in marine organisms. Harmful Marine Algal Blooms, pp.475-480, 1995.

B. Petton, F. Pernet, R. Robert, and P. Boudry, Temperature influence on pathogen transmission and subsequent mortalities in juvenile Pacific oysters Crassostrea gigas, Aquac. Environ. Inter, vol.3, pp.257-273, 2013.
URL : https://hal.archives-ouvertes.fr/hal-00946757

É. Pousse, J. Flye-sainte-marie, M. Alunno-bruscia, H. Hégaret, and F. Jean, Sources of paralytic shellfish toxin accumulation variability in the pacific oyster crassostrea gigas, Toxicon, 2017.
URL : https://hal.archives-ouvertes.fr/hal-02324600

S. Pouvreau, Y. Bourles, S. Lefebvre, A. Gangnery, and M. Alunno-bruscia, Application of a dynamic energy budget model to the Pacific oyster, Crassostrea gigas, reared under various environmental conditions, J. Sea Res, vol.56, issue.2, pp.156-167, 2006.
URL : https://hal.archives-ouvertes.fr/hal-00148899

S. M. Renaud, L. Thinh, G. Lambrinidis, and D. L. Parry, Effect of temperature on growth, chemical composition and fatty acid composition of tropical Australian microalgae grown in batch cultures, Aquaculture, vol.211, issue.1, pp.195-214, 2002.

C. S. Reynolds, The ecology of phytoplankton, 2006.

J. Rolland, W. Medhioub, A. Vergnes, C. Abi-khalil, V. Savar et al., A feedback mechanism to control apoptosis occurs in the digestive gland of the oyster Crassostrea gigas exposed to the paralytic shellfish toxins producer Alexandrium catenella, Mar. Drugs, vol.12, issue.9, pp.5035-5054, 2014.

M. Rosa, J. E. Ward, A. Frink, and S. E. Shumway, Effects of surface properties on particle capture by two species of suspension-feeding bivalve molluscs, Am. Malacol. Bull, vol.35, issue.2, pp.181-188, 2017.

M. Rosa, J. E. Ward, B. A. Holohan, S. E. Shumway, and G. H. Wikfors, Physicochemical surface properties of microalgae and their combined effects on particle selection by suspension-feeding bivalve molluscs, J. Exp. Mar. Biol. Ecol, vol.486, pp.59-68, 2017.

M. Rosa, J. E. Ward, S. E. Shumway, G. H. Wikfors, E. Pales-espinosa et al., Effects of particle surface properties on feeding selectivity in the eastern oyster Crassostrea virginica and the blue mussel Mytilus edulis, J. Exp. Mar. Biol. Ecol, vol.446, pp.320-327, 2013.

S. Saraiva, J. Van-der-meer, S. Kooijman, and T. Sousa, Modelling feeding processes in bivalves : a mechanistic approach, Ecol. Model, vol.222, issue.3, pp.514-523, 2011.

S. E. Shumway, A review of the effects of algal blooms on shellfish and aquaculture, J. World Aquac. Soc, vol.21, issue.2, pp.65-104, 1990.

S. E. Shumway and T. L. Cucci, The effects of the toxic dinoflagellate Protogonyaulax tamarensis on the feeding and behaviour of bivalve molluscs, Aquat. Toxicol, vol.10, issue.1, pp.9-27, 1987.

W. Silvert and A. Cembella, Dynamic modelling of phycotoxin kinetics in the blue mussel, Mytilus edulis, with implications for other marine invertebrates, Can. J. of Fish. Aquat. Sci, vol.52, issue.3, pp.521-531, 1995.

P. Soletchnik, D. Razet, and P. Geairon, Ecophysiology of maturation and spawning in oyster (Crassostrea gigas) : metabolic (respiration) and feeding (clearance and absorption rates) responses at different maturation stages, Oceanogr. Lit. Rev, vol.12, issue.44, p.1545, 1997.

Y. Thomas, S. Pouvreau, M. Alunno-bruscia, L. Barillé, F. Gohin et al., Global change and climate-driven invasion of the Pacific oyster (Crassostrea gigas) along European coasts : a bioenergetics modelling approach, J. Biogeogr, vol.26, 2016.
URL : https://hal.archives-ouvertes.fr/hal-02530683

R. Van-haren, H. Schepers, and S. Kooijman, Dynamic energy budgets affect kinetics of xenobiotics in the marine mussel Mytilus edulis, Chemosphere, vol.29, issue.2, pp.163-189, 1994.

P. Visciano, M. Schirone, M. Berti, A. Milandri, R. Tofalo et al., Marine biotoxins : occurrence, toxicity, regulatory limits and reference methods, Front. Microbiol, vol.7, 2016.

J. Ward, J. Levinton, S. Shumway, and T. Cucci, Particle sorting in bivalves : in vivo determination of the pallial organs of selection, Mar. Biol, vol.131, issue.2, pp.283-292, 1998.

J. E. Ward and S. E. Shumway, Separating the grain from the chaff : particle selection in suspension-and deposit-feeding bivalves, J. Exp. Mar. Biol. Ecol, vol.300, issue.1, pp.83-130, 2004.

T. Yamamoto, K. J. Flynn, and H. Takayama, Application of a two-compartment one-toxin model to predict the toxin accumulation in Pacific oysters in Hiroshima Bay, Japan. Fish. Sci, vol.69, issue.5, pp.944-950, 2003.

T. Yan, M. Zhou, M. Fu, Y. Wang, R. Yu et al., Inhibition of egg hatching success and larvae survival of the scallop, Chlamys farreri, associated with exposure to cells and cell fragments of the dinoflagellate Alexandrium tamarense, Toxicon, vol.39, issue.8, pp.1239-1244, 2001.

T. Yasumoto and M. Murata, Marine toxins, Chem. Rev, vol.93, issue.5, pp.1897-1909, 1993.

S. A. Kooijman, Dynamic energy and mass budgets in biological systems, 2000.

S. A. Kooijman, Dynamic energy budget theory for metabolic organisation, 2010.

Y. Thomas, S. Pouvreau, M. Alunno-bruscia, L. Barillé, F. Gohin et al., Global change and climate-driven invasion of the Pacific oyster (Crassostrea gigas) along European coasts : a bioenergetics modelling approach, J. Biogeogr, vol.43, issue.3, pp.568-579, 2016.
URL : https://hal.archives-ouvertes.fr/hal-02530683

H. W. Van-der-veer, J. F. Cardoso, and J. Van-der-meer, The estimation of deb parameters for various Northeast Atlantic bivalve species, J. Sea Res, vol.56, issue.2, pp.107-124, 2006.

J. Whyte, J. Englar, and B. Carswell, Biochemical composition and energy reserves in Crassostrea gigas exposed to different levels of nutrition, Aquaculture, vol.90, issue.2, pp.157-172, 1990.